Site last updated on: May 2, 2017

Contact Dr. Beran: mjberan@yahoo.com

Journal Publications (student names in boldface) - Click title to see the abstract. Please contact us for reprints.

Beran, M. J. (2017). To err is (not only) human: Fallibility as a window into primate cognition.  Comparative Cognition & Behavior Reviews, 12, 57-81. Click here for a PDF file of this paper.

Parrish, A. E., James, B. J., & Beran, M. J. (2017). Exploring whether nonhuman primates show a bias to overestimate dense quantities. Journal of Comparative Psychology, 131, 59-68.

Agrillo, C., Parrish, A. E., & Beran, M. J. (2016). How illusory is the Solitaire
illusion? Assessing the degree of misperception of numerosity in adult humans.
Frontiers in Psychology, 7,1663. doi: 10.3389/fpsyg.2016.01663. Click here for a PDF file of this paper.

Beran, M. J. (2016). "Zeroing” in on mathematics in the monkey brain. Learning & Behavior, 44, 4-6.

Beran, M. J., Hopper, L. M., de Waal, F. B. M., Brosnan, S. F., & Sayers, K. (2016). Chimpanzees, cooking, and comparative psychology. Learning & Behavior, 44, 103-108.

Beran, M. J., Hopper, L. M., de Waal, F. B. M., Sayers, K., & Brosnan, S. F. (2016). Chimpanzee food preferences, associative learning, and the origins of cooking. Learning & Behavior, 44, 118-121.

Beran, M. J., James, B. T., Whitham, W., & Parrish, A. E. (2016). Chimpanzees can point to smaller amounts of food to accumulate larger amounts but they still fail the reverse-reward contingency task. Journal of Experimental Psychology: Animal Learning and Cognition, 42, 347-358.

Beran, M. J., Menzel, C. R., Parrish, A. E., Perdue, B. M., Sayers, K., Smith, J. D., & Washburn, J. D. (2016). Primate cognition: Attention, episodic memory, prospective memory, self-control, and metacognition as examples of cognitive control in nonhuman primates. WIREs Cognitive Science.  doi: 10.1002/wcs.1397

Beran, M. J., & Parrish, A. E. (2016). Capuchin monkeys (Cebus apella) treat small and large numbers of items similarly during a relative quantity judgment task. Psychonomic Bulletin & Review, 23, 1206-1213.

Beran, M. J., Perdue, B. M., Church, B. A., & Smith, J. D. (2016). Capuchin monkeys (Cebus apella) modulate their use of an uncertainty response depending on risk. Journal of Experimental Psychology: Animal Learning and Cognition, 42, 32-43.

Beran, M. J., Perdue, B. M., Rossettie, M. S., James, B. T., Whitham, W., Walker, B., Futch, S. E., & Parrish, A. E. (2016). Self-control assessments of capuchin monkeys with the rotating tray task and the accumulation task.  Behavioural Processes, 129, 68-79. 

Beran, M. J., Rossettie, M. S., & Parrish, A. E. (2016). Trading up: Chimpanzees (Pan troglodytes) show self-control through their exchange behavior. Animal Cognition, 19, 109-121.

Parrish, A. E., Agrillo, C., Perdue, B. M., & Beran, M. J. (2016). The elusive illusion: Do children (Homo sapiens) and capuchin monkeys (Cebus apella) see the Solitaire illusion? Journal of Experimental Child Psychology, 142, 83-95.

Parrish, A. E., Emerson, I. D., Rossettie, M. S., & Beran, M. J. (2016). Testing the ego-depletion hypothesis among capuchin monkeys: Does glucose boost self-control? Behavioral Sciences, 6, 16. Click here for full-text PDF file.

Agrillo, C., Gori, S., & Beran, M. J. (2015). Do rhesus monkeys (Macaca mulatta) perceive illusory motion? Animal Cognition, 18, 895-910.

Beran, M. J. (2015). Chimpanzee cognitive control. Current Directions in Psychological Science, 24, 352-357.

Beran, M. J. (2015). The comparative science of "self-control": What are we talking about? Frontiers in Psychology, 6, Article 51. Click here for full-text PDF file.

Beran, M. J. (2015). Animal memory: Chimpanzees anticipate what comes next in short movies. Current Biology, 25, R829-R830.

Beran, M. J., & Heimbauer, L. A. (2015). A longitudinal assessment of vocabulary retention in symbol-competent chimpanzees (Pan troglodytes).  PLoS ONE, 10, e0118408. Click here for full-text PDF file.

Beran, M. J., James, B. T., Futch, S. E., & Parrish, A. E. (2015). A comparative psychologist by any other name…is still a comparative psychologist. Commentary: A crisis in comparative psychology: Where have all the undergraduates gone? Frontiers in Psychology, 6, 1928. Click here for full-text PDF file.

Beran, M. J., Parrish, A. E., Futch, S. E., Evans, T. A., & Perdue, B. M. (2015).  Looking ahead? Computerized maze task performance by chimpanzees (Pan troglodytes), rhesus monkeys (Macaca mulatta), capuchin monkeys (Cebus apella), and human children (Homo sapiens). Journal of Comparative Psychology, 129, 160-173.

Beran, M. J., Perdue, B. M., & Evans, T. A. (2015). Prospective memory in nonhuman primates. Japanese Journal of Animal Psychology, 65, 23-33.

Beran, M. J., Perdue, B. M., Futch, S. E., Smith, J. D., Evans, T. A., & Parrish, A. E. (2015). Go when you know: Chimpanzees’ confidence movements reflect their responses in a computerized memory task. Cognition, 142, 236-246.

Parrish, A. E., Brosnan, S. F., & Beran, M. J. (2015). Do you see what I see? A comparative investigation of the Delboeuf illusion in humans (Homo sapiens), rhesus monkeys (Macaca mulatta) and capuchin monkeys (Cebus apella). Journal of Experimental Psychology: Animal Learning and Cognition, 41, 395-405.

Parrish, A. E., Brosnan, S. F., & Beran, M. J. (2015). Capuchin monkeys alternate play and reward in a dual computerized task. Animal Behavior and Cognition, 2, 334-347.

Parrish, A. E., Evans, T. A., & Beran, M. J. (2015). Rhesus macaques (Macaca mulatta) exhibit the decoy effect in a perceptual discrimination task.  Attention, Perception, & Psychophysics, 77, 1715-1725.

Parrish, A. E., Evans, T. A., & Beran, M. J. (2015). Defining value through quantity and quality - Chimpanzees (Pan troglodytes) undervalue food quantities when items are broken.  Behavioural Processes, 111, 118-126.

Perdue, B. M., Bramlett, J. L., Evans, T. A., & Beran, M. J. (2015). Waiting for what comes later: Capuchin monkeys show self-control even for nonvisible delayed rewards. Animal Cognition, 18, 1105-1112.

Perdue, B. M., Church, B. A., Smith, J. D., & Beran, M. J. (2015). Exploring potential mechanisms underlying the lack of uncertainty monitoring in capuchin monkeys. International Journal of Comparative Psychology, Article 28. Click here for full-text PDF file.

Sayers, K., Evans, T. A., Menzel, E., Smith, J. D., & Beran, M. J. (2015). The misbehaviour of a metacognitive monkey. Behaviour, 152, 727-756.

Agrillo, C., Parrish, A. E., & Beran, M. J. (2014). Do primates see the Solitaire illusion differently? A comparative assessment of Humans (Homo sapiens), chimpanzees (Pan troglodytes), rhesus monkeys (Macaca mulatta) and capuchin monkeys (Cebus apella).  Journal of Comparative Psychology, 128, 402-413.

Agrillo, C., Parrish, A. E., & Beran, M. J. (2014). Do rhesus monkeys (Macaca mulatta) perceive the Zöllner illusion?  Psychonomic Bulletin & Review, 21, 986-994.

Beran, M. J., Evans, T. A., Paglieri, F., McIntyre, J. M., Addessi, E., & Hopkins, W. D. (2014). Chimpanzees (Pan troglodytes) can wait, when they choose to: A study with the hybrid delay task.  Animal Cognition, 17, 197-205.

Beran, M. J., Parrish, A. E., Perdue, B. M., & Washburn, D. A. (2014). Comparative cognition: Past, present, and future.  International Journal of Comparative Psychology, 27, 3-30. Click here for full-text PDF file.

Beran, M. J., Perdue, B. M., & Smith, J. D. (2014). What are my chances? Closing the gap in uncertainty monitoring between rhesus monkeys (Macaca mulatta) and capuchin monkeys (Cebus apella). Journal of Experimental Psychology: Animal Learning and Cognition, 40, 303-316.

Beran, M. J., & Smith, J. D. (2014). The uncertainty response in animal-metacognition researchers.  Journal of Comparative Psychology, 128, 155-159.

Evans, T. A., & Beran, M. J. (2014). Monkeys wait to begin a computer task when waiting makes their responses more effective. Animal Behavior and Cognition, 1, 36-50.

Evans, T. A., Perdue, B. M., Parrish, A. E., & Beran, M. J. (2014). The relationship between event-based prospective memory and ongoing task performance in chimpanzees (Pan troglodytes).  PLoS ONE, 9, e112015.

Evans, T. A., Perdue, B. M., Parrish, A. E., & Beran, M. J. (2014). Working and waiting for better rewards: Self-control in two monkey species (Cebus apella and Macaca mulatta).  Behavioural Processes, 103, 236-242.

Garland, A., Beran, M. J., McIntyre, J. M., & Low, J. (2014). Relative quantity judgments between discrete spatial arrays by chimpanzees (Pan trogloydytes) and New Zealand robins (Petroica longipesJournal of Comparative Psychology, 128, 307-317.

Parrish, A. E., & Beran, M. J. (2014). When less is more: Like humans, chimpanzees (Pan troglodytes) misperceive food amounts based on plate size. Animal Cognition, 17, 427-434.

Parrish, A. E., & Beran, M. J. (2014). Chimpanzees sometimes see fuller as better: Judgments of food quantities based on container size and fullness. Behavioural Processes, 103, 184-191.

Parrish, A. E., Brosnan, S. F., Wilson, B. J., & Beran, M. J. (2014).  Differential responding by rhesus monkeys (Macaca mulatta) and humans (Homo sapiens) to variable outcomes in the Assurance game. Animal Behavior and Cognition, 1, 215-229.

Parrish, A. E., Perdue, B. M., Stromberg, E. E., Bania, A. E., Evans, T. A., & Beran, M. J. (2014). Delay of gratification by orangutans (Pongo pygmaeus) in the accumulation task.  Journal of Comparative Psychology, 128, 209-214.

Perdue, B. M., Beran, M. J., Williamson, R. A., Gonsiorowski, A., Evans, T. A. (2014). Prospective memory in children and chimpanzees.  Animal Cognition, 17, 287-295.

Perdue, B. M., Evans, T. A., Washburn, D. A., Rumbaugh, D. M., & Beran, M. J. (2014). Do monkeys choose to choose?  Learning & Behavior, 42, 164-175.

Smith, J. D., Couchman, J. J., & Beran, M. J. (2014). Animal metacognition: A tale of two comparative psychologies.  Journal of Comparative Psychology, 128, 115-131.  

Smith, J. D., Couchman, J. J., & Beran, M. J. (2014). Animal metacognition: A tale of two comparative psychologies – Reply to Commentaries.  Journal of Comparative Psychology, 128, 140-142.

Vonk, J., Torgerson-White, L., McGuire, M., Thueme, M., Thomas, J., & Beran, M. J. (2014). Quantity estimation and comparison in Western Lowland Gorillas (Gorilla gorilla gorilla). Animal Cognition, 17, 755-765.

Zakrzewski, A. C., Perdue, B. M., Beran, M. J., Church, B. A., & Smith, J. D. (2014). Cashing out: The decisional flexibility of uncertainty responses in rhesus macaques (Macaca mulatta) and humans (Homo sapiens). Journal of Experimental Psychology: Animal Learning and Cognition, 40, 490-501.

Addessi, E., Paglieri, F., Beran, M. Evans, T. Macchitella, L., De Petrillo, F., & Focaroli, V. (2013).  Delay choice vs. delay maintenance: Different measures of delayed gratification in capuchin monkeys (Cebus apella).  Journal of Comparative Psychology, 127, 392-398.

Beran, M. J., McIntyre, J. M., Garland, A., & Evans, T. A. (2013).  What counts for “counting”? Chimpanzees (Pan troglodytes) respond appropriately to relevant and irrelevant information in a quantity judgment task.  Animal Behaviour, 85, 987-993.

Beran, M. J., & Parrish, A. E. (2013). Visual nesting of stimuli affects rhesus monkeys’ (Macaca mulatta) quantity judgments in a bisection task. Attention, Perception, & Psychophysics, 75, 1243-1251.

Beran, M. J., Smith, J. D., & Perdue, B. M. (2013).  Language-trained chimpanzees name what they have seen, but look first at what they have not seen.  Psychological Science, 24, 660-666.

Brosnan, S. F., Beran, M. J., Parrish, A. E., Price, S. A., & Wilson, B. J. (2013). Comparative approaches to studying strategy: Towards an evolutionary account of primate decision-making.  Evolutionary Psychology, 11, 606-627.

Klein, E. D., Evans, T. A., Schultz, N. B., & Beran, M. J. (2013). Learning how to “make a deal”: Human and monkey performance when repeatedly faced with the Monty Hall Dilemma. Journal of Comparative Psychology, 127, 103-108.

Paglieri, F., Focaroli, V., Bramlett, J., Tierno, V., McIntyre, J., Addessi, E., Evans, T. A., & Beran, M. J. (2013).  The hybrid delay task: Can capuchin monkeys (Cebus apella) sustain a delay after an initial choice to do so? Behavioural Processes, 94, 45-54.

Parrish, A. E., Perdue, B. M., Evans, T. A., & Beran, M. J. (2013). Chimpanzees (Pan troglodytes) transfer token repeatedly with a partner to accumulate rewards in a self-control task. Animal Cognition, 16, 627-636.

Smith, J. D., Coutinho, M. V. C., Church, B., & Beran, M. J. (2013). Executive-attentional uncertainty responses by rhesus monkeys (Macaca mulatta).  Journal of Experimental Psychology: General, 142, 458-475.

Smith, J. D., Flemming, T. M., Boomer, J., Beran, M. J., & Church, B. A. (2013). Fading perceptual resemblance: A path for rhesus macaques (Macaca mulatta) to conceptual matching?  Cognition, 129, 598-614.

Beran, M. J. (2012). Did you ever hear the one about the horse that could count?  Frontiers in Comparative Psychology, Article 257. Click here for full-text PDF file.

Beran, M. J. (2012). Quantity judgments of auditory and visual stimuli by chimpanzees (Pan troglodytes). Journal of Experimental Psychology: Animal Behavior Processes, 38, 23-29.

Beran, M. J., Decker, S., Schwartz, A., & Smith, J. D. (2012). Uncertainty monitoring by young children in a computerized task.  Scientifica, 2012, Article 692890. Click here for full-text PDF file.

Beran, M. J., & Evans, T. A. (2012).  Language-trained chimpanzees (Pan troglodytes) delay gratification by choosing token exchange over immediate reward consumption.  American Journal of Primatology, 74, 864-870.

Beran, M. J., Evans, T. A., Klein, E. D., & Einstein, G. O. (2012). Rhesus monkeys (Macaca mulatta) and capuchin monkeys (Cebus apella) remember future responses in a computerized task. Journal of Experimental Psychology: Animal Behavior Processes, 38, 233-243.

Beran, M. J., Owens, K., Phillips, H. A., & Evans, T. A. (2012). Humans and monkeys show similar skill in estimating uncertain outcomes. Psychonomic Bulletin & Review, 19, 357-362.

Beran, M. J., & Parrish, A. E. (2012). Sequential responding and planning in capuchin monkeys (Cebus apella).  Animal Cognition, 15, 1085-1094.

Beran, M. J., Perdue, B. M., Bramlett, J. L., Menzel, C. R., & Evans, T. A. (2012). Prospective memory in a language-trained chimpanzee (Pan troglodytes). Learning and Motivation, 43, 192-199. 

Beran, M. J., Perdue, B. M., Parrish, A. E., & Evans, T. A. (2012). Do social conditions affect capuchin monkeys’ (Cebus apella) choices in a quantity judgment task? Frontiers in Comparative Psychology, 3, Article 492.  Click here for full-text PDF file.

Bramlett, J. L., Perdue, B. M., Evans, T. A., & Beran, M. J. (2012). Capuchin monkeys (Cebus apella) let lesser rewards pass them by to get better rewards. Animal Cognition, 15, 963-969. 

Brosnan, S. F., Wilson, B. J., & Beran, M. J. (2012). Old World monkeys are more similar to humans than New World monkeys when playing a coordination game.  Proceedings of the Royal Society of London, 279, 522-1530.

Couchman, J. J., Beran, M. J., Coutinho, M. V. C., Boomer, J., Zakrzewski, A., Church, B. J., & Smith, J. D. (2012). Do actions speak louder than words? A comparative perspective on implicit vs. explicit metacognition and theory of mind. British Journal of Developmental Psychology, 30, 210-221.

Evans, T. A., & Beran, M. J. (2012).  Monkeys exhibit prospective memory in a computerized task.  Cognition, 125, 131-140.

Evans, T. A., Beran, M. J., Paglieri, F., Addessi, E. (2012). Delaying gratification for food and tokens in capuchin monkeys (Cebus apella) and chimpanzees (Pan troglodytes): When quantity is salient, symbolic stimuli do not improve performance. Animal Cognition, 15, 539-548.

Evans, T. A., Perdue, B. M., Parrish, A. E., Menzel, E., Brosnan, S. F., & Beran, M. J. (2012). How is chimpanzee self-control influenced by social setting? Scientifica, Article ID 654094. Click here for full-text PDF file.

Heimbauer, L. A., Conway, C. M., Christiansen, M. H., Beran, M. J., & Owren, M. J. (2012).  A serial reaction time (SRT) task with symmetrical joystick responding for nonhuman primates.  Behavior Research Methods, 44, 733-741.

Perdue, B. M., Talbot, C. F., Stone, A., & Beran, M. J. (2012).  Putting the elephant back in the herd: Elephant relative quantity judgments match those of other species.  Animal Cognition, 15, 955-961.

Smith, J. D., Berg, M. E., Cook, R. G., Murphy, M. S., Crossley, M. J., Boomer, J., Spiering, B., Beran, M. J., Church, B. A., Ashby, F. G., & Grace, R. C. (2012).  Implicit and explicit categorization: A tale of four species.  Neuroscience and Biobehavioral Reviews, 36, 2355-2369.

Smith, J. D., Couchman, J. J., & Beran, M. J. (2012). The highs and lows of theoretical interpretation in animal-metacognition research. Philosophical Transactions of the Royal Society B, 367, 1297-1309.

Smith, J. D., Crossley, M. J., Boomer, J., Church, B. A., Beran, M. J., & Ashby, F. G. (2012). Implicit and explicit category learning by capuchin monkeys (Cebus apella). Journal of Comparative Psychology, 126, 294-304.

Vonk, J., & Beran, M. J. (2012). Bears “count” too: Quantity estimation and comparison in black bears (Ursus americanus).  Animal Behaviour, 84, 231-238.

Beran, M. J. (2011). Chimpanzees (Pan troglodytes) show the isolation effect during serial list recognition memory tests. Animal Cognition, 14, 637-645.

Beran, M. J., Decker, S., Schwartz, A., & Schultz, N. (2011). Monkeys (Macaca mulatta and Cebus apella) and human adults and children (Homo sapiens) enumerate and compare subsets of moving stimuli based on numerosity. Frontiers in Comparative Psychology, 2, Article 61. Click here for full-text PDF file.

Beran, M. J., Evans, T. A., & Hoyle, D. (2011). Numerical judgments by chimpanzees (Pan troglodytes) in a token economy. Journal of Experimental Psychology: Animal Behavior Processes, 37, 165-174.

Beran, M. J., Johnson-Pynn, J. S., & Ready, C. (2011). Comparing children’s (Homo sapiens) and chimpanzees’ (Pan troglodytes) quantity judgments of sequentially presented sets of items. Current Zoology, 57, 419-428.

Beran, M. J., & Smith, J. D. (2011). Information seeking by rhesus monkeys (Macaca mulatta) and capuchin monkeys (Cebus apella). Cognition, 120, 90-105.

Brosnan, S. F., Parrish, A. E., Beran, M. J., Flemming, T., Heimbauer, L., Talbot, C., Lambeth, S. P., Shapiro, S. J., Wilson, B. J. (2011). Responses to the Assurance game in monkeys, apes, and humans using equivalent procedures. Proceedings of the National Academy of Sciences, 108, 3442-3447.

Flemming, T. M., Thompson, R. K. R., Beran, M. J., & Washburn, D. A. (2011).  Analogical reasoning and the differential outcome effect: Transitory bridging of the conceptual gap for rhesus monkeys (Macaca mulatta).  Journal of Experimental Psychology: Animal Behavior Processes, 37, 353-360.

Heimbauer, L. A., Beran, M. J., & Owren, M. J. (2011).  A chimpanzee recognizes synthetic speech with significantly reduced acoustic cues to phonetic content.  Current Biology, 21, 1210-1214. 

Klein, E. D., Evans, T. A., & Beran, M. J. (2011). An investigation of prospective and retrospective coding in capuchin monkeys and rhesus monkeys. Zeitschrift für Psychologie / Journal of Psychology, 219, 85-91.

Beran, M. J. (2010). Chimpanzees (Pan troglodytes) accurately compare poured liquid quantities. Animal Cognition, 13, 641-649.

Beran, M. J. (2010). Use of exclusion by a chimpanzee (Pan troglodytes) during speech perception and auditory-visual matching-to-sample. Behavioural Processes, 83, 287-291.

Couchman, J. J., Coutinho, M. V. C., Beran, M. J., Smith, J. D. (2010). Beyond stimulus cues and reinforcement signals: A new approach to animal metacognition. Journal of Comparative Psychology, 124, 356-368.

Evans, T. A., Beran, M. J., & Addessi, E. (2010). Can nonhuman primates use tokens to represent and sum quantities? Journal of Comparative Psychology, 124, 369-380. 

Harris, E. H., Gulledge, J. P., Beran, M. J., & Washburn, D. A. (2010). What do Arabic numerals mean to macaques? Journal of Experimental Psychology: Animal Behavior Processes, 36, 66-76.

Smith, J. D., Beran, M. J., Crossley, M., Boomer, J., & Ashby, F. G. (2010). Implicit and explicit category learning by macaques (Macaca mulatta) and humans (Homo sapiens). Journal of Experimental Psychology: Animal Behavior Processes, 36, 54-65.

Smith, J. D., Redford, J. S., Beran, M. J., & Washburn, D. A. (2010).  Rhesus monkeys (Macaca mulatta) adaptively monitor uncertainty while multi-tasking. Animal Cognition, 13, 93-101. 

Washburn, D. A., Gulledge, J. P., Beran, M. J., & Smith, J. D. (2010). With his memory magnetically erased, a monkey knows he is uncertain. Biology Letters, 6, 160-162.

Beran, M. J. (2009). Chimpanzees as natural accountants.  Human Evolution, 24, 183-196. Click here for full-text PDF file.

Beran, M. J., & Evans, T. A. (2009). Delay of gratification by chimpanzees (Pan troglodytes) in working and waiting situations. Behavioural Processes, 80, 117-121.

Beran, M. J., Evans, T. A., & Harris, E. H. (2009). When in doubt, chimpanzees rely on estimates of past reward amounts.  Proceedings of the Royal Society B, 276, 309-314.

Beran, M. J., Evans, T. A., & Ratliff, C. L. (2009). Perception of food amounts by chimpanzees (Pan troglodytes): The role of magnitude, contiguity, and wholeness. Journal of Experimental Psychology: Animal Behavior Processes, 35, 516-524.

Beran, M. J., Ratliff, C. L., & Evans, T. A. (2009). Natural choice in chimpanzees (Pan troglodytes): Perceptual and temporal effects on selective value. Learning and Motivation, 40, 186-196.  

Beran, M. J., Smith, J. D., Coutinho, M. V. C., Couchman, J. J., & Boomer, J. (2009). The psychological organization of “uncertainty” responses and “middle” responses: A dissociation in capuchin monkeys (Cebus apella). Journal of Experimental Psychology: Animal Behavior Processes, 35, 371-381.

Brosnan, S. F., & Beran, M. J. (2009). Trade between conspecifics in chimpanzees (Pan troglodytes). Journal of Comparative Psychology, 123, 181-194.

Evans, T. A. , Beran, M. J., Harris, E. H., & Rice, D. (2009).  Quantity judgments of sequentially presented food items by capuchin monkeys (Cebus apella). Animal Cognition, 12, 97-105.

Hoffman, M. L., Beran, M. J., & Washburn, D. A. (2009).  Memory for 'what,' 'where,' and 'when' information in rhesus monkeys (Macaca mulatta).  Journal of Experimental Psychology: Animal Behavior Processes, 35, 143-152.

Smith, J. D., Beran, M. J., Couchman, J. J., Coutinho, M. V. C., & Boomer, J. (2009). Animal metacognition: Problems and prospects. Comparative Cognition and Behavior Reviews, 4, 33-46.

Beran, M. J. (2008). Monkeys (Macaca mulatta and Cebus apella) track, enumerate, and compare multiple sets of moving items. Journal of Experimental Psychology: Animal Behavior Processes, 34, 63-74.

Beran, M. J. (2008).  Capuchin monkeys (Cebus apella) succeed in a test of quantity conservation. Animal Cognition, 11, 109-116.

Beran, M. J. (2008).  The evolutionary and developmental foundations of mathematics.  PLoS Biology, 6, e19.

Beran, M. J., Evans, T. A., & Harris, E. H. (2008). Perception of food amount by chimpanzees based on the number, size, contour length, and visibility of items. Animal Behaviour, 75, 1793-1802.

Beran, M. J., Evans, T. A., Leighty, K. A., Harris, E. H., & Rice, D. (2008).  Summation and quantity judgments of sequentially presented sets by capuchin monkeys (Cebus apella). American Journal of Primatology, 70, 191-194.

Beran, M. J., Harris, E. H., Evans, T. A., Klein, E. D., Chan, B., Flemming, T. M., & Washburn, D. A. (2008). Ordinal judgments of symbolic stimuli by capuchin monkeys (Cebus apella) and rhesus monkeys (Macaca mulatta): The effects of differential and nondifferential reward.  Journal of Comparative Psychology, 122, 52-61. 

Beran, M. J., Johnson-Pynn, J. S., & Ready, C. (2008).  Quantity representation in children and rhesus monkeys: Linear versus logarithmic scales.  Journal of Experimental Child Psychology, 100, 225-233. 

Beran, M. J., Klein, E. D., Evans, T. A., Chan, B., Flemming, T. M., Harris, E. H., Washburn, D. A., & Rumbaugh, D. M. (2008). Discrimination reversal learning in capuchin monkeys (Cebus apella).  Psychological Record, 58, 3-14.

Brosnan, S. F., Grady, M. F., Lambeth, S. P., Schapiro, S. J., & Beran, M. J. (2008). Chimpanzee autarky. PloS ONE, 3, e1518.

Evans, T. A., Beran, M. J., Chan, B., Klein, E. D., & Menzel, C. R. (2008).  An efficient computerized testing method for the capuchin monkey (Cebus apella): Adaptation of the LRC-CTS to a socially housed nonhuman primate species.  Behavior Research Methods, 40, 590-596. 

Flemming, T. M., Beran, M. J., Thompson, R. K. R., Kleider, H. M., & Washburn, D. A. (2008).  What meaning means for same and different: Analogical reasoning in humans (Homo sapiens), chimpanzees (Pan troglodytes), and rhesus monkeys (Macaca mulatta).  Journal of Comparative Psychology, 122, 176-185.

Smith, J. D., Beran, M. J., Couchman, J. J., & Coutinho, M. V. C. (2008). The comparative study of metacognition: Sharper paradigms, safer inferences. Psychonomic Bulletin and Review, 15, 679-691.

Beran, M. J. (2007). Rhesus monkeys (Macaca mulatta) enumerate large and small sequentially presented sets of items using analog numerical representations. Journal of Experimental Psychology: Animal Behavior Processes, 33, 42-54.

Beran, M. J. (2007). Rhesus monkeys (Macaca mulatta) succeed on a computerized test designed to assess conservation of discrete quantity. Animal Cognition, 10, 37-45.

Beran, M. J., Rumbaugh, D. M., & Washburn, D. A. (2007). Rhesus monkeys (Macaca mulatta) maintain learning set despite second-order stimulus-response spatial discontiguity. Psychological Record, 57, 9-22.

Beran, M. J., Washburn, D. A., & Rumbaugh, D. M. (2007).  The Stroop Effect in color-naming of color-word lexigrams by a chimpanzee.  Journal of General Psychology, 134, 217-228. 

Evans, T. A., & Beran, M. J. (2007). Chimpanzees use self-distraction to cope with impulsivity. Biology Letters, 3, 599-602. 

Evans, T. A., & Beran, M. J. (2007). Delay of gratification and delay maintenance by rhesus macaques (Macaca mulatta). Journal of General Psychology, 134, 199-216.

Flemming, T. M., Beran, M. J., & Washburn, D. A. (2007). Disconnect in concept learning by rhesus monkeys: Judgment of relations and relations-between-relations. Journal of Experimental Psychology: Animal Behavior Processes, 33, 55-63.

Harris, E. H., Beran, M. J., & Washburn, D. A. (2007). Ordinal list integration for symbolic, arbitrary, and analog stimuli by rhesus macaques (Macaca mulatta). Journal of General Psychology, 134, 183-197.

Harris, E. H., Washburn, D. A., Beran, M. J., & Sevcik, R. A. (2007). Rhesus monkeys (Macaca mulatta) select Arabic numerals or visible quantities corresponding to a number of sequentially completed maze trials. Learning and Behavior, 35, 53-59.

Beran, M. J. (2006).  Quantity perception by adult humans (Homo sapiens), chimpanzees (Pan troglodytes), and rhesus macaques (Macaca mulatta) as a function of stimulus organization. International Journal of Comparative Psychology, 19, 386-397. 

Beran, M. J., & Evans, T. A. (2006). Maintenance of delay of gratification by four chimpanzees (Pan troglodytes): The effects of delayed reward visibility, experimenter presence, and extended delay intervals. Behavioural Processes, 73, 315-324.

Beran, M. J., Smith, J. D., Redford, J. S., & Washburn, D. A. (2006). Rhesus macaques (Macaca mulatta) monitor uncertainty during numerosity judgments. Journal of Experimental Psychology: Animal Behavior Processes, 32, 111-119.

Beran, M. J., Taglialatela, L. B., Flemming, T. M., James, F. M., & Washburn, D. A. (2006). Nonverbal estimation during numerosity judgments by adult humans. Quarterly Journal of Experimental Psychology, 59, 2065-2082.

Hoffman, M. L., & Beran, M. J. (2006). Chimpanzees (Pan troglodytes) remember the location of a hidden food item after altering their orientation to a spatial array. Journal of Comparative Psychology, 120, 389-393.

Smith, J. D., Beran, M. J., Redford, J. S., & Washburn, D. A. (2006). Dissociating uncertainty responses and reinforcement signals in the comparative study of uncertainty monitoring. Journal of Experimental Psychology: General, 135, 282-297.

Beran, M. J., Beran, M. M., Harris, E. H., & Washburn, D. A. (2005).  Ordinal judgments and summation of nonvisible sets of food items by two chimpanzees (Pan troglodytes) and a rhesus macaque (Macaca mulatta).  Journal of Experimental Psychology: Animal Behavior Processes, 31, 351-362. 

Beran, M. J., Beran, M. M., & Menzel, C. R. (2005). Chimpanzees (Pan troglodytes) use markers to monitor the movement of a hidden food item. Primates, 46, 255-259.

Beran, M. J., Beran, M. M., & Menzel, C. R. (2005). Spatial memory and monitoring of hidden items through spatial displacements by chimpanzees (Pan troglodytes). Journal of Comparative Psychology, 119, 14-22.

Beran, M. J. (2004). Long-term retention of the differential values of Arabic numerals by chimpanzees (Pan troglodytes). Animal Cognition, 7, 86-92.

Beran, M. J. (2004). Chimpanzees (Pan troglodytes) respond to nonvisible sets after one-by-one addition and removal of items. Journal of Comparative Psychology, 118, 25-36.

Beran, M. J., & Beran, M. M. (2004). Chimpanzees remember the results of one-by-one addition of food items to sets over extended time periods. Psychological Science, 15, 94-99.

Beran, M. J., Pate, J. L., Washburn, D. A., & Rumbaugh, D. M. (2004). Sequential responding and planning in chimpanzees (Pan troglodytes) and rhesus macaques (Macaca mulatta). Journal of Experimental Psychology: Animal Behavior Processes, 30, 203-212.

Beran, M. J., & Washburn, D. A. (2002). Chimpanzee responding during matching to sample: Control by exclusion. Journal of the Experimental Analysis of Behavior, 78, 497-508.

Beran, M. J. (2002). Maintenance of self-imposed delay of gratification by four chimpanzees (Pan troglodytes) and an orangutan (Pongo pygmaeus). Journal of General Psychology, 129, 49-66.

Beran, M. J. (2001). Summation and numerousness judgments of sequentially presented sets of items by chimpanzees (Pan troglodytes). Journal of Comparative Psychology, 155, 181-191.

Beran, M. J. (2001). Do chimpanzees have expectations about reward presentation following correct performance on computerized cognitive testing? Psychological Record, 51, 173-183.

Beran, M. J., & Rumbaugh, D. M (2001). "Constructive" enumeration by chimpanzees (Pan troglodytes) on a computerized task. Animal Cognition, 4, 81-89.

Beran, M. J., & Minahan, M. F. (2000). Monitoring spatial transpositions by bonobos (Pan paniscus) and chimpanzees (Pan troglodytes). International Journal of Comparative Psychology, 13, 1-15.

Beran, M. J., Pate, J. L., Richardson, W. K., & Rumbaugh, D. M. (2000). A chimpanzee's (Pan troglodytes) long-term retention of lexigrams. Animal Learning and Behavior, 28, 201-207.

Beran, M. J., Savage-Rumbaugh, E. S., Pate, J. L., & Rumbaugh, D. M. (1999). Delay of gratification in chimpanzees (Pan troglodytes). Developmental Psychobiology, 34, 119-127.

Beran, M. J., Rumbaugh, D. M., & Savage-Rumbaugh, E. S. (1998). Chimpanzee (Pan troglodytes) counting in a computerized testing paradigm. Psychological Record, 48, 3-20.

Beran, M. J., Savage-Rumbaugh, E. S., Brakke, K. E., Kelley, J. W., & Rumbaugh, D. M. (1998). Symbol comprehension and learning: A "vocabulary" test of three chimpanzees (Pan troglodytes). Evolution of Communication, 2, 171-188.